Skip to content


B.S., Massachusetts Institute of Technology, 1952. Ph.D., Massachusetts Institute of Technology, 1955 (D. F. Waugh). Postdoctoral: Naval Medical Research Institute, Bethesda, Maryland, 1956 (M. F. Morales). Honors and Awards: NSF Postdoctoral Fellow, 1952-55; Public Health Service Senior Research Fellow (Career Development Award) 1959-67; John S. Guggenheim Memorial Foundation Fellow, 1973-74; Elected Member, National Academy of Sciences, 1978; Elected Fellow, American Academy of Arts and Sciences, 1979; Merck Award, American Society of Biochemistry and Molecular Biology, 2002; Elected Member, American Philosophical Society, 2004; Editor, Journal Molecular Biology, 1986-94; Editorial Board, Trends in Biochemical Sciences, 1986 - present; Member, Advisory Committee to the Director, NIH, 1987 - 92; American Cancer Society Research Professor of Chemistry, 1989 - present. At Oregon since 1967.